diff --git a/gcc/cp/module.cc b/gcc/cp/module.cc
index 999ff3faafc..2c2f9a9a8cb 100644
--- a/gcc/cp/module.cc
+++ b/gcc/cp/module.cc
@@ -10034,15 +10034,11 @@ trees_out::tpl_parms_fini (tree tmpl, unsigned tpl_levels)
       tree vec = TREE_VALUE (parms);
 
       tree_node (TREE_TYPE (vec));
-      tree dflt = error_mark_node;
       for (unsigned ix = TREE_VEC_LENGTH (vec); ix--;)
 	{
 	  tree parm = TREE_VEC_ELT (vec, ix);
-	  if (dflt)
-	    {
-	      dflt = TREE_PURPOSE (parm);
-	      tree_node (dflt);
-	    }
+	  tree dflt = TREE_PURPOSE (parm);
+	  tree_node (dflt);
 
 	  if (streaming_p ())
 	    {
@@ -10072,19 +10068,15 @@ trees_in::tpl_parms_fini (tree tmpl, unsigned tpl_levels)
        tpl_levels--; parms = TREE_CHAIN (parms))
     {
       tree vec = TREE_VALUE (parms);
-      tree dflt = error_mark_node;
 
       TREE_TYPE (vec) = tree_node ();
       for (unsigned ix = TREE_VEC_LENGTH (vec); ix--;)
 	{
 	  tree parm = TREE_VEC_ELT (vec, ix);
-	  if (dflt)
-	    {
-	      dflt = tree_node ();
-	      if (get_overrun ())
-		return false;
-	      TREE_PURPOSE (parm) = dflt;
-	    }
+	  tree dflt = tree_node ();
+	  if (get_overrun ())
+	    return false;
+	  TREE_PURPOSE (parm) = dflt;
 
 	  tree decl = TREE_VALUE (parm);
 	  if (TREE_CODE (decl) == TEMPLATE_DECL)
diff --git a/gcc/testsuite/g++.dg/modules/pr105045_a.C b/gcc/testsuite/g++.dg/modules/pr105045_a.C
new file mode 100644
index 00000000000..597f9294185
--- /dev/null
+++ b/gcc/testsuite/g++.dg/modules/pr105045_a.C
@@ -0,0 +1,7 @@
+// PR c++/105045
+// { dg-additional-options -fmodules-ts }
+// { dg-module-cmi pr105045 }
+
+export module pr105045;
+
+export template<int T=0, class U> void f(U) { }
diff --git a/gcc/testsuite/g++.dg/modules/pr105045_b.C b/gcc/testsuite/g++.dg/modules/pr105045_b.C
new file mode 100644
index 00000000000..77c94d4c473
--- /dev/null
+++ b/gcc/testsuite/g++.dg/modules/pr105045_b.C
@@ -0,0 +1,6 @@
+// PR c++/105045
+// { dg-additional-options -fmodules-ts }
+
+import pr105045;
+
+int main() { f(0); }